The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.

Systematics

A considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or incertae sedis.

1. Subfamily Berothimerobiinae

2. Berothimerobius

3. Subfamily Berothinae

4. Asadeteva

5. Barrowiella

6. Berlekrumyia

7. Berotha

8. †Elektroberotha Eocene (Lutetian); Baltic amber, northern Europe

9. Isoscelipteron

10. Lekrugeria

11. Lomamyia

12. Nodalla

13. Podallea

14. Quasispermophorella

15. Spermophorella

16. Spiroberotha

17. Stenobiella

18. †Xenoberotha Green River Formation, United States, Eocene

19. Subfamily Cyrenoberothinae

20. †Aggregataberotha (Cenomanian, Burmese amber, Myanmar)

21. †Araripeberotha Cretaceous (Aptian); Crato Formation, Brazil

22. †Caririberotha Cretaceous (Aptian); Crato Formation, Brazil

23. Cyrenoberotha

24. Manselliberotha

25. †Microberotha (Eocene (Ypresian), Hat Creek amber, Canada)

26. Ormiscocerus

27. †Protoberotha (Cenomanian, Burmese amber, Myanmar)

28. †Sibelliberotha (Barremian, Lebanese amber)

29. Speleoberotha

30. Subfamily †Mesithoninae

31. †Mesithone Callovian/Oxfordian; Karabastau Formation, Kazakhstan; Aptian; Zaza Formation, Turga Formation, Russia

32. †Oloberotha Aptian; Yixian Formation, China

33. Subfamily Nosybinae

34. †Banoberotha Cretaceous (Barremian); Lebanese amber, Lebanon

35. Naizema

36. Nosybus

37. Tanzanberotha

38. Subfamily Nyrminae

39. Nyrma

40. Subfamily Protobiellinae

41. Austroberothella

42. Protobiella

43. Subfamily Trichomatinae

44. Trichoberotha

45. Trichoma

46. Subfamily Incertae sedis

47. †Ansoberotha Cenomanian; Burmese amber, Myanmar

48. †Berothone Callovian/Oxfordian; Karabastau Formation, Kazakhstan

49. †Cantabroberotha Cretaceous (Albian); Spanish amber, Spain

50. †Cornoberotha Cenomanian; Burmese amber, Myanmar

51. †Dasyberotha Cenomanian; Burmese amber, Myanmar

52. †Epimesoberotha Cretaceous (Berriasian); Lulworth Formation, United Kingdom

53. †Ethiroberotha Cenomanian; Burmese amber, Myanmar

54. †Haploberotha Cenomanian; Burmese amber, Myanmar

55. †Iceloberotha Cenomanian; Burmese amber, Myanmar

56. †Jersiberotha Cenomanian; Burmese amber, Myanmar; Cretaceous (Turonian); New Jersey amber, United States

57. †Krokhathone Jurassic (Callovian/Oxfordian); Karabastau Formation, Kazakhstan

58. †Maculaberotha Cenomanian; Burmese amber, Myanmar

59. †Magniberotha Cenomanian; Burmese amber, Myanmar

60. †Nascimberotha Turonian; New Jersey amber, United States

61. †Plesiorobius Cretaceous (Campanian); Canadian amber, Canada; Campanian; Taymyr amber, Russia; Cenomanian, unknown formation, Russia

62. †Proberotha Lutetian, Baltic Amber, northern Europe

63. †Osmyloberotha Cenomanian; Burmese amber, Myanmar

64. †Sinosmylites Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, Ulaan-Ereg Formation, Mongolia, Late Jurassic (Tithonian)

65. †Systenoberotha Cenomanian; Burmese amber, Myanmar

66. †Telistoberotha Cenomanian; Burmese amber, Myanmar

67. †Xiaoberotha Cenomanian; Burmese amber, Myanmar

Formerly included taxa

Subfamily Paraberothinae (now included in Rhachiberothidae)

• †Acanthoberotha Cenomanian; Burmese amber, Myanmar

• †Alboberotha Cretaceous (Albian); Charentense amber, France

• †Astioberotha Cenomanian; Burmese amber, Myanmar

• †Chimerhachiberotha Barremian; Lebanese amber, Lebanon

• †Creagroparaberotha Cenomanian; Burmese amber, Myanmar

• †Eorhachiberotha Cenomanian; Burmese amber, Myanmar

• †Kujiberotha Cretaceous (Santonian); Kuji amber, Japan

• †Micromantispa Cenomanian; Burmese amber, Myanmar

• †Paraberotha Barremian; Lebanese amber, Lebanon

• †Pseudosisyra Aptian; Zaza Formation, Russia

• †Raptorapax Barremian; Lebanese amber, Lebanon

• †Retinoberotha Cenomanian; Bezonnais amber, France,

• †Rhachibermissa Turonian; New Jersey amber, United States

• †Scoloberotha Cenomanian; Burmese amber, Myanmar

• †Spinoberotha Cenomanian; Burmese amber, Myanmar

• †Stygioberotha Cenomanian; Burmese amber, Myanmar

• †Uranoberotha Cenomanian; Burmese amber, Myanmar

References