↑ arvau

Wikipedia / Urocoptidae

Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea. <ref name="WoRMS">MolluscaBase eds. (2026). MolluscaBase. Urocoptidae Pilsbry, 1898 (1868). Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=932609 on 2026-07-02

Taxonomy

2005 taxonomy

The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

2008 taxonomy

Uit de Weerd (2008) moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.

The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea. There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.

2012 taxonomy

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (Epirobia, Propilsbrya, Pectinistemma, Gyrocion, Prionoloplax), that were previously placed within Urocoptidae. Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.

Genera

Genera in the family Urocoptidae include:

  1. Brachypodellinae

  2. Apoma H. Beck, 1837

  3. Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae

  4. Cyclauchen Pilsbry, 1930

  5. Geoscala Pilsbry & Vanatta, 1898

  6. Ischnostrophina Pilsbry, 1933

  7. Mychostoma Albers, 1850

  8. Simplicervix Pilsbry, 1903

  9. Urocoptinae

  10. Acrocoptis C. de la Torre & Bartsch, 1972

  11. Allocoptis Thompson & Franz, 1976

  12. Amphicosmia Pilsbry & Vanatta, 1898

  13. Amphistemma C. de la Torre & Bartsch, 1972

  14. Anafecoptis C. de la Torre & Bartsch, 1972

  15. Angulicervix Pilsbry, 1933

  16. Anoma Albers, 1850

  17. Arangia Pilsbry & Vanatta, 1898

  18. Archegocoptis Pilsbry, 1903

  19. Autocoptis Pilsbry, 1902

  20. Bactrocoptis Pilsbry, 1902

  21. Badiofaux Pilsbry, 1941

  22. Bialasmus C. de la Torre & Bartsch, 1972

  23. Brachypodella L. Pfeiffer, 1840 (as Cylindrella)

  24. Bostrichocentrum Strebel, 1879 : synonym of Holospira E. von Martens, 1860

  25. Brevipedella Pilsbry, 1903

  26. Callocoptis C. de la Torre & Bartsch, 1972

  27. Callonia Crosse & P. Fischer, 1870

  28. Capillacea C. de la Torre & Bartsch, 1972

  29. Carcinostemma C. de la Torre & Jaume, 1972

  30. Centralia de la Torre & Bartsch, 1972

  31. Cochlodinella Pilsbry & Vanatta, 1898

  32. Esochara Pilsbry & Vanatta, 1898

  33. Fibricutis Pilsbry, 1903

  34. Geminicoptis C. de la Torre & Bartsch, 1972

  35. Gongylostoma Albers, 1850

  36. Gyraxis Pilsbry, 1903

  37. Heterocoptis C. de la Torre & Bartsch, 1972

  38. Idiostemma Pilsbry & Vanatta, 1898

  39. Insulaceramus Clench, 1967

  40. Johaniceramus C. de la Torre & Bartsch, 1972

  41. Levistemma C. de la Torre & Bartsch, 1972

  42. Liocallonia Pilsbry, 1902

  43. Liparotes Pilsbry, 1903

  44. Maceo Pilsbry & Vanatta, 1898

  45. Macroceramus Guilding, 1828

  46. Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae

  47. Nesocoptis Pilsbry, 1941

  48. Nodulia C. de la Torre & Bartsch, 1972

  49. Organocoptis C. de la Torre & Bartsch, 1972

  50. Orienticoptis C. de la Torre & Bartsch, 1972

  51. Paracallonia Pilsbry, 1903

  52. Pfeiffericoptis C. de la Torre & Bartsch, 1972

  53. Pineria Poey, 1854

  54. Planostemma C. de la Torre & Bartsch, 1972

  55. Pleurostemma C. de la Torre & Bartsch, 1972

  56. Poecilocoptis Pilsbry, 1941

  57. Poeycoptis C. de la Torre & Bartsch, 1972

  58. Pseudopineria Aguayo, 1938

  59. Pycnoptychia Pilsbry & Vanatta, 1898

  60. Sagracoptis C. de la Torre & Bartsch, 1972

  61. Sectilumen Pilsbry & Vanatta, 1898

  62. Spiroceramus Pilsbry & Vanatta, 1898

  63. Spirostemma Pilsbry & Vanatta, 1898

  64. Steatocoptis Pilsbry, 1941

  65. Strophina Mörch, 1852

  66. Teneria C. de la Torre & Bartsch, 1972

  67. Tenuistemma C. de la Torre & Bartsch, 1972

  68. Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae

  69. Tomelasmus Pilsbry & Vanatta, 1898

  70. Torrecoptis Bartsch, 1943

  71. Trilamellaxis C. de la Torre & Bartsch, 1972

  72. Uncinicoptis C. de la Torre & Bartsch, 1972

  73. Urocoptis Beck, 1837 - type genus of the family Urocoptidae

  74. Urocoptidae incertae sedis (temporary name)

  75. Habeas Simone, 2013

References

Further reading

  • ; 7 figs.

  • Carlos de la Torre. 1912. New Cuban Urocoptis of the U. cinerea group. The Nautilus, volume 26, number 4, pages 55-58, plate 6.

  • Pilsbry H. A. & Henderson J. B. jr. (February 1913). Two new Cuban Urocoptidae. The Nautilus, volume 26, number 10, pages 109-110.

  • Miguel L. Jaume & Alfredo de la Torre. 1976. Los Urocoptidae de Cuba (Mollusca-Pulmonata). Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp.

  • Richardson C. L. 1991. Urocoptidae: catalogue of species. Tryonia 22: 1–245.

  • Tryon G. W. (1903) Manual of Conchology. Second series: Pulmonata. Volume 15. Urocoptidae. Continued by H. A. Pilsbry.