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Wikipedia / Urocyclidae

Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea.

Some species within this family make and use love darts before mating.

Distribution

The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.

Taxonomy

The family Urocyclidae consists of 3 subfamilies:

  • subfamily Rhysotininae Schileyko, 2002

  • subfamily Sheldoniinae Connolly, 1925 (1912) - synonyms: Peltatinae Godwin-Austen, 1912; Trochonanininae Connolly, 1912; Trochozonitinae Iredale, 1914; Ledoulxiinae Pilsbry, 1919; Gymnarionidae Von Mol, 1970; Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002

subfamily Urocyclinae Simroth, 1889

Genera

The following genera are recognised in the family Urocyclidae:

  1. unsorted

  2. † Koruella Pickford, 2009

  3. Subfamily Rhysotininae

  4. Rhysotina Ancey, 1887

  5. Subfamily Sheldoniinae

  6. Acantharion Binder & Tillier, 1985

  7. Africarion Godwin-Austen, 1883

  8. Amatarion Van Mol, 1970

  9. Angustivestis Pilsbry, 1919

  10. Aspidelus Morelet, 1884

  11. Aspidotomium Degner, 1932

  12. Belonarion Pilsbry, 1919

  13. Bloyetia Bourguignat, 1890

  14. Camerunarion Van Mol, 1970

  15. Carinazingis van Bruggen & de Winter, 1990

  16. Centrafricarion Van Mol, 1970

  17. Chlamydarion Van Mol, 1968

  18. Colparion Laidlaw, 1938

  19. Degneria Verdcourt, 1956

  20. Entagaricus Pilsbry, 1919

  21. Estria Poirier, 1887

  22. Falloonella Preston, 1914

  23. Fuellebornia Verdcourt, 1998

  24. Granularion Germain, 1912

  25. Gymnarion Pilsbry, 1919

  26. †Hamya Bourguignat, 1885

  27. Kerkophorus Godwin-Austen, 1912

  28. Lacrimarion Connolly, 1939

  29. Malagarion Tillier, 1979

  30. Mesafricarion Pilsbry, 1919

  31. Microkerkus Godwin-Austen, 1912

  32. Montanobloyetia Verdcourt, 1961

  33. Morrumbalia Verdcourt, 1998

  34. Paragranularion Van Mol, 1970

  35. Percivalia Preston, 1914

  36. Plicatonanina Verdcourt, 1961

  37. Principicochlea D. Holyoak & G. Holyoak, 2020

  38. Principitrochoidea D. Holyoak & G. Holyoak, 2020

  39. Ptilototheca Herbert, 2016

  40. Rhopalogonium Degner, 1932

  41. Selatodryas Herbert, 2017

  42. Senegalarion Van Mol, 1970

  43. Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae

  44. Sjostedtina Verdcourt, 1961

  45. Sylvarion Van Mol, 1970

  46. Thapsia Albers, 1860

  47. Thielarion Van Mol, 1970

  48. Thomithapsia D. Holyoak & G. Holyoak, 2020

  49. Thomitrochoidea D. Holyoak & G. Holyoak, 2020

  50. Tresia Van Goethem, 1975

  51. Trochonanina Mousson, 1869

  52. Trochozonites Pfeffer, 1883

  53. Tropidocochlion Verdcourt, 1998

  54. Verdcourtia Van Mol, 1970

  55. Verrucarion Van Mol, 1970

  56. Zingis E. von Martens, 1878

  57. Zonitarion Pfeffer, 1883

  58. Subfamily Urocyclinae

Tribe Dendrolimacini

  • Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini

Tribe Leptichnini

  • Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini

Tribe Upembellini

tribe Urocyclini

Cladogram

The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:

References