Wikipedia / Characidae
Wikipedia / Characidae
Characidae, the characids, is a family of freshwater subtropical and tropical fish belonging to the order Characiformes. They are found throughout much of Central and South America, including such major waterways as the Amazon and Orinoco Rivers. These fish vary in length; many are less than .
The name "characins" is a historical one, but scientists today tend to prefer "characids" to reflect their status as a, by and large, monophyletic group (at family rank). This family includes some of the first characiforms to be described to science, such as Charax and Tetragonopterus, and thus lend their name to the order, as well as to common names such as "characin" and "tetra".
Past taxonomic treatments had a much more expansive definition of the family, including numerous South American fish families such as the piranhas and dorados, as well as the African alestids. Following multiple taxonomic revisions, this was eventually restricted to just the American "tetra" type characins by the 2010s. However, even this definition of Characidae was found to obscure much of the evolutionary diversity within the group, and in 2024 the "tetra" families Acestrorhamphidae and Stevardiidae were split out of the Characidae, leaving it with just five subfamilies.
Classification
Taxonomy
The following classification is based on Eschmeyer's Catalog of Fishes (2025): Family Characidae
Subfamily Aphyocharacinae C. H. Eigenmann, 1909
Amazonichthys Esguícero & Mendonça, 2023
Aphyocharacidium Géry, 1960
Aphyocharax Günther, 1868
Cyanogaster Mattox, Britz, Toledo-Piza & Marinho, 2013
Leptagoniates Boulenger, 1887
Paragoniates Steindachner, 1876
Phenagoniates C. H. Eigenmann & C. B. Wilson, 1914
Prionobrama Fowler, 1913
Xenagoniates Myers, 1942
Subfamily Cheirodontinae C. H. Eigenmann, 1915
Acinocheirodon Malabarba & Weitzman, 1999
Amblystilbe Fowler, 1940
Aphyocheirodon C. H. Eigenmann, 1915
Cheirodontops Schultz, 1944
Compsura C. H. Eigenmann, 1915
Ctenocheirodon Malabarba & Jerep, 2012
Heterocheirodon Malabarba, 1998
Holoshesthes C. H. Eigenmann, 1903
Kolpotocheirodon Malabarba & Weitzman, 2000
Macropsobrycon C. H. Eigenmann, 1915
Nanocheirodon Malabarba, 1998
Odontostilbe Cope, 1870
Prodontocharax C. H. Eigenmann & Pearson, 1924
Protocheirodon Vari, Melo & Oliveira, 2016
Pseudocheirodon Meek & Hildebrand, 1916
Saccoderma Schultz, 1944
Serrapinnus Malabarba 1998
Subfamily Exodontinae Fowler, 1958
Bryconexodon Géry, 1980
Roeboexodon Géry, 1959
Subfamily Tetragonopterinae Gill, 1858
Tetragonopterus Cuvier, 1816
Subfamily Characinae Latreille, 1825
Acanthocharax C. H. Eigenmann, 1912
Acestrocephalus C. H. Eigenmann, 1910
Atopomesus Myers, 1927
Cynopotamus Valenciennes, 1850
Galeocharax Fowler, 1910
Microschemobrycon C. H. Eigenmann, 1915
Phenacogaster C. H. Eigenmann, 1907
Roeboides Günther 1864
Former members
This family has undergone a large amount of systematic and taxonomic change. More recent revision has moved many former members of the family into their own related but distinct families – the pencilfishes of the genus Nannostomus are a typical example, having now been moved into the Lebiasinidae, the assorted predatory species belonging to Hoplias and Hoplerythrinus have now been moved into the Erythrinidae, and the sabre-toothed fishes of the genus Hydrolycus have been moved into the Cynodontidae. The former subfamily Alestiinae was promoted to family level (Alestiidae) and the subfamilies Crenuchinae and Characidiinae were moved to the family Crenuchidae.
Other fish families that were formerly classified as members of the Characidae, but which were moved into separate families of their own during recent taxonomic revisions (after 1994) include Acestrorhynchidae, Anostomidae, Chilodidae, Citharinidae, Ctenoluciidae, Curimatidae, Distichodontidae, Gasteropelecidae, Hemiodontidae, Hepsetidae, Parodontidae, Prochilodontidae, Serrasalmidae, and Triportheidae. In 2024, the families Stevardiidae and Acestrorhamphidae, containing a high proportion of the famous ornamental aquarium tetras, were also split out of the family, in addition to the small family Spintherobolidae.
References
de Lucena, Carlos Alberto Santos (2003): New characid fish, Hyphessobrycon scutulatus, from the rio Teles Pires drainage, upper rio Tapajós system (Ostariophysi: Characiformes: Characidae). Neotropical Ichthyology 1(2): 93–96. PDF fulltext
Géry, Jacques (1977): Characoids of the World.
Nelson, Joseph S. (2006): Fishes of the World. John Wiley & Sons, Inc.