Wikipedia / Polygalaceae
Wikipedia / Polygalaceae
The Polygalaceae or the milkwort family are made up of flowering plants in the order Fabales. They have a near-cosmopolitan range, with about 27 genera and ca. 900 known species of herbs, shrubs and trees. Over half of the species are in one genus, Polygala, the milkworts.
The family was first described in1809 by Johann Hoffmansegg and Johann Link. In1896, Robert Chodat split it into three tribes. A fourth tribe was split off from the tribe Polygaleae in1992. Under the Cronquist classification system, Polygalaceae were treated in a separate order of their own, Polygalales. Currently, according to the Angiosperm Phylogeny Group, the family belongs in Fabales.
Description
Polygalaceae are annual or perennial herbs, shrubs, trees or lianas. Its zygomorphic, hermaphrodite, bisexual flowers have three to five petals and five sepals. Its leaves are usually alternate, but may be opposite, fascicled, or verticillate. Each flower usually contains eight stamens, though this may range from three to ten. They are usually in two series. The fruits of each plant can be a capsule, samara, or drupe.
Tribes and genera
The Polygalaceae comprise the following genera, with tribes based on various sources.
Carpolobieae Eriksen
Atroxima Stapf
Carpolobia G. Don
Moutabeae Chodat
Barnhartia Gleason
Diclidanthera Mart.
Eriandra P. Royen & Steenis
Moutabea Aubl.
Polygaleae Chodat
Acanthocladus Klotzsch ex Hassk.
Asemeia Raf.
Badiera DC.
Bredemeyera Willd.
Caamembeca J.F.B. Pastore
Chamaebuxus (synonym Polygaloides Haller)
Comesperma Labill.
Epirixanthes Blume
Gymnospora (Chodat) J.F.B.Pastore
Hebecarpa (Chodat) J.R.Abbott
Heterosamara Kuntze
Hualania Phil.
Monnina Ruiz & Pav.
Monrosia Grondona
Muraltia DC.
Phlebotaenia Griseb.
Polygala L.
Rhamphopetalum J.F.B.Pastore & M.Mota
Rhinotropis (S.F. Blake) J.R. Abbott
Salomonia Lour.
Securidaca L.
Senega (DC.) Spach
Xanthophylleae Chodat
Xanthophyllum Roxb.
Fossils
†Paleosecuridaca Pigg, Kathleen B., M.L. DeVore & M.F. Wojc. 2008
Systematics
Modern molecular phylogenetics suggest the following relationships:
References